Late Successional Mixed Conifer Forests

Coast Range Ecoregion
Late Successional Mixed Conifer Forest within Oswald West State Park. Photo Credit: David Patte, USFWS

Late successional mixed conifer forests provide a multi-layered tree canopy, including large-diameter trees, shade-tolerant tree species in the understory, and a high volume of dead wood, such as snags and logs.

Ecoregions

Late successional mixed conifer forests are a Key Habitat in the Blue Mountains, Coast Range, East Cascades, Klamath Mountains, and West Cascades ecoregions.

Characteristics

Late successional mixed conifer forests are defined by plant species composition, overstory tree age and size, and the forest structure. While a range of tree ages are present in late successional forests, the predominant stand age is over 150 years. They include characteristics such as a multi-layered tree canopy, shade-tolerant tree species growing in the understory, large-diameter trees, and a high volume of dead wood, such as snags and logs. These characteristics can take hundreds of years without stand-replacing disturbance to develop. Historically, fire was the major natural disturbance in all but the wettest areas. Depending on local conditions, fires in the Coast Range and West Cascades conifer forests were of moderate- to high-severity, with fire return intervals averaging 100 to more than 400 years. These stand replacing events were interspersed with periodic low severity understory burns every 15 to 30 years. This historical fire regime created a complex mosaic of stand structures across the landscape.

Ecoregional Characteristics

Blue Mountains

A mixture of conifer species is found throughout many forest sites in the Blue Mountains ecoregion. Mixed conifer forests can be divided into two subtypes based on temperature and moisture conditions. Douglas-fir and grand fir are the primary late successional tree species in the warmer drier climates of this ecoregion. Ponderosa pine and western larch may also be present. The cool mixed conifer type is indicated by the addition of more moisture-demanding and cold-tolerant species, such as subalpine fir and Engelmann spruce, at upper elevations or along streams where cold water-drainage and deep frost eliminate some species. The understory in this ecoregion generally includes huckleberry, serviceberry, oceanspray, snowberry, wild ginger, goldthread, starflower, bead lily, and oak fern.

Coast Range

Late successional mixed conifer forests in the Coast Range are generally dominated by two types of conifer trees: Sitka spruce and Douglas-fir. Sitka spruce forests occur within a narrow fog- and salt-influenced strip along the coast and extend into some valleys. Soils tend to be deep, acidic, and well-drained. Sitka spruce dominates the overstory, but western hemlock, western redcedar, Douglas-fir, big leaf maple, and red alder may be present. The lush understory has salmonberry, vine maple, salal, evergreen huckleberry, sword fern, deer fern, and a high diversity of mosses and lichens. Inland, Douglas-fir forests dominate. The understory of Douglas-fir forests includes shrub and forb species, such as vine maple, salal, sword fern, Cascade Oregon grape, western rhododendron, huckleberries, twinflower, vanilla leaf, and oxalis. Due to high precipitation in both Sitka spruce and Douglas-fir forests, fires are infrequent but do occur during hot, dry, east wind conditions after prolonged drought. When fires do occur, they are likely to be high severity, stand-replacing events.  Other disturbances include small-scale windthrow events and floods driven by atmospheric river storms.

East Cascades

Late successional mixed conifer forests span the eastern slopes of the Cascade Mountains. This habitat contains a wide variety of tree species with Douglas-fir, grand fir, and western hemlock as the most common forest tree species that co-dominate most overstories. Several other conifers may also be present, including western redcedar, western white pine, western larch, ponderosa pine, and lodgepole pine. Undergrowth vegetation in the East Cascades ecoregion includes vine maple, Oregon grape, huckleberry, oxalis, thimbleberry, manzanita, ceanothus, and twinflower. Many sites once dominated by Douglas-fir and ponderosa pine (formerly maintained by wildfire) may now be dominated by grand fir (a fire sensitive, shade-tolerant species).

Klamath Mountains

Late successional mixed conifer forests in the Klamath Mountains ecoregion are characterized by high tree diversity. Douglas-fir is usually dominant. Depending on site characteristics, other canopy trees may include white fir, sugar pine, ponderosa pine, and incense cedar. Port-Orford cedar occurs on moist sites, such as riparian areas. Jeffrey pine and knobcone pine occur on serpentine soils. Broadleaf trees, such as tanoak, canyon live oak, golden chinquapin, big leaf and vine maple, and Pacific madrone, may occur in the subcanopy. A range of understory communities may be present, including those mostly dominated by shrubs, forbs, or grasses, or may be relatively open. However, with an increase in frequency of droughts, high instances of Douglas-fir mortality in the Klamath Mountains is quickly shifting the composition of these forests, creating conditions that result in catastrophic wildfires.

West Cascades

Late successional mixed conifer forests are found scattered throughout the West Cascades Ecoregion. While Douglas -fir dominates these forests, western hemlock is almost always co-dominant and usually dominates the understory. In the absence of stand replacing disturbance, Douglas-fir forests eventually convert to western hemlock. Other common trees include grand fir and western redcedar in the northern portion of the ecoregion, or incense cedar, sugar pine, white fir, and western redcedar in the southern portion of the ecoregion. The understory has shrub and forb species, such as vine maple, salal, sword fern, Cascade Oregon grape, western rhododendron, huckleberries, twinflower, vanilla leaf, and oxalis.

Conservation Overview

Since the 1850s, both timber harvest and large-scale fires have replaced many of the late successional forests in Oregon with younger forests. Many of the remaining late successional forests occur in a patchwork with the younger forests that are managed with shorter rotations to generate timber products. While a mosaic of forest age classes contributes to landscape-scale diversity, many species associated with late-successional forests require large patches of older or mature forests to survive and may be sensitive to changes in the forest seral stage. Late successional mixed conifer forests are particularly important for wildlife, mosses, and lichens. Depending on the ecoregion, Species of Greatest Conservation Need associated with late successional conifer forests include ringtailfisherPacific martenred tree voleMarbled MurreletNorthern Spotted OwlOregon slender salamander, and many others.

The Northwest Forest Plan (NWFP) is a comprehensive natural resource planning effort that includes all or part of the Siuslaw, Rogue River-Siskiyou. Mt. Hood, Willamette, Deschutes, Umpqua, and Fremont Winema National Forests in Oregon. The NWFP identifies conservation priorities for species affected by loss and fragmentation of large patches of late successional forests, assessing over 1,000 species. Late Successional Reserves established under the NWFP are intended to ensure enough high-quality habitat to sustain identified species. However, many of the federal lands that are designated as Late Successional Reserves do not include forests at the late successional stage, while others are relatively small “checkerboards” of forests embedded in a matrix of private industrial timber lands, particularly in the Coast Range and Klamath Mountains.

Late successional mixed conifer forests are generally declining on federal lands in the NWFP area, though at a slower rate than before the plan’s implementation. From 1994 until 2020, there was an increase in the acres of late successional stage forest in the NWFP area. However, in 2020, wildland fires driven by strong east winds in the Mt. Hood, Willamette, Umpqua, and Rogue Siskiyou National Forests destroyed many acres of late successional forest. The overall goal of the NWFP is to achieve desired conditions for these forests over decades, a goal that remains feasible but requires sustained effort to manage disturbances like wildfire in fire-prone areas. Particular attention is needed for the fire-prone portions of the NWFP area, where large wildfires have set back forest conditions from desired outcomes.

The US Forest Service’s Wildfire Crisis Strategy Implementation Plan and Oregon Department of Forestry’s 20 Year Landscape Resiliency Strategy are examples of federal and state efforts to address uncharacteristic wildfire in Oregon’s forests. These plans address historical fire suppression, exclusion of cultural burning practices, and the impacts of recent catastrophic and uncharacteristic wildfires by recommending a variety of active management techniques for forests to increase forest resiliency to wildland fire.

Oregon’s state forests are managed to support a healthy balance of timber production, clean water, recreation, and wildlife habitat. The Oregon Department of Forestry (ODF) oversees the Northwest and Southwest State Forest Management Plans, which guide how Board of Forestry lands are managed. These management plans include conservation strategies for both upland forest and riparian habitats and support a wide range of SGCN – including both terrestrial and aquatic species. ODF is currently developing the Western Oregon State Forest Management Plan and the Western Oregon State Forest Habitat Conservation Plan (HCP), which when approved will replace the Northwest and Southwest Oregon State Forest Management Plans and cover about 64,000 acres of ODF managed lands west of the Cascades.

The Oregon Department of State Lands (DSL) manages’ the Elliot State Research Forest, Oregon’s first state-designated research forest. DSL developed a Habitat Conservation Plan (HCP) for the Elliot to protect late-successional forest habitat, which is critical to SGCN such as Marbled Murrelet and Northern Spotted Owl. The Elliot HCP balances forest research and limited management activities with long-term habitat conservation goals.

The Private Forest Accord (PFA) is a compromise agreement made between representatives from Oregon’s timber industry, the Oregon Small Woodlands Association, and prominent conservation and fishing organizations, to modify portions of Oregon’s forest practice laws and regulations in a way that expands protections for fish and amphibians. The changes to the Oregon Forest Practices Act are aimed at avoiding and minimizing the effects that timber harvests and other forest management activities on private forestlands have on these species and the aquatic habitats they depend on. The PFA included new standards for stream classification and protection or stream buffers, steep slopes, roads, and culverts, as well as a grant program to fund riparian and stream habitat restoration projects. These standards may also help to restore some late successional mixed conifer forests located in riparian areas adjacent to streams.

In the southwest Cascades, white firs in overstocked stands often have a high prevalence  of disease, creating a buildup of fuels and putting large ponderosa pines at risk for severe wildfire. Removing these patches of dead, dying, and diseased trees will help create more early seral openings for the benefit of deer and elk and provide a greater mosaic of habitats across the landscape.

Limiting Factors and Recommended Approaches

Limiting Factor: Loss of Structural Habitat Elements

Where historical stands were perpetuated for 200 to more than 1,000 years, commercial forestlands are now commonly harvested every 60 years or less, which limits the maintenance and future recruitment of late-successional characteristics. In addition, the number of large-diameter snags and logs, which contribute to understory structure, has been reduced over time through wildfire and timber harvest.

Recommended Approach

Develop programs, incentives, and market-based approaches to encourage longer rotations and strategically located areas where large-diameter trees predominate. Where feasible, maintain structural elements, such as large-diameter tall trees, snags, and logs. Create snags from green trees or high-cut stumps where maintaining snags is not feasible or where snag management goals are not being met. Maintain old forest stand structures on private industrial forest lands. Provide technical assistance to landowners to leave large-diameter downed wood, green trees, or snags in the upland portion of harvested forests, as well as along riparian areas, to provide benefits for a diversity of wildlife and fish. Follow Oregon Forest Practices Act rules, which help to maintain a diversity of structural components on the landscape.

Limiting Factor: Loss of Late Successional Stand Size and Connectivity

Late successional forest stands have been greatly reduced in size and connectivity, particularly at lower elevations. This can impact species that are highly adapted to late successional conditions, require large tracts of intact habitats, and/or species that have limited ability to move over long distances to find new suitable areas. It also allows edge species to compete with those adapted to extensive interior forest habitat.

Recommended Approach

Maintain existing plans to protect and develop late successional habitat. Use active management to accelerate development of late successional structural characteristics in key areas to expand existing patches into larger areas; these will provide greater blocks of habitat for species with large area requirements or those that require interior forest habitat and are vulnerable to “edge effects”. Continue to carefully plan forest practices to maintain connectivity (KCIBarriers to Animal Movement), particularly when species vulnerable to fragmentation are present. ODFW has mapped Priority Wildlife Connectivity Areas (PWCAs) to provide information on places across the landscape with the highest overall value for facilitating wildlife movement.

Seek opportunities to coordinate management of public and private lands (e.g., All-Lands Approach) whenever possible to address conservation needs. Use voluntary conservation tools, such as financial incentives and forest certification to achieve conservation goals on private lands. Work to maintain a diversity of forest types and ages to support wildlife habitat connectivity and ecosystem function at a landscape scale.

HABITAT CHANGE TRENDS ANALYSIS

Late successional mixed conifer forest habitats have been greatly reduced in size and connectivity by timber harvest. Following disturbance, such as timber harvest, regenerating conifer forests often succeed to deciduous or mixed deciduous-conifer forests without active management (e.g. replanting with Douglas-fir), especially on the west side of the state. These deciduous forests are primarily dominated by red alder and bigleaf maple. To investigate the transition from late successional mixed conifer forests to deciduous forest, the Institute of Natural Resources (INR) analyzed the change in total area of west-side deciduous and mixed conifer-deciduous forests between 1851 and 2016. The analysis showed a massive 243% increase in deciduous forests between 1851 and 1998, likely reflecting a shift towards earlier successional forests following logging.

Resources for more information

Northwest Forest Plan

Elliott State Research Forest Habitat Conservation Plan

Oregon Department of State Lands

Oregon Department of Forestry

Western Oregon State Forest Habitat Conservation Plan

Status and Trends of Late Successional and Old Growth Forests

Oregon Private Forest Accord

REFERENCES

Brunner, R. and E. Gaines. 2025. Oregon Vegetation Change 1851-2023. Trends analysis conducted for Oregon Department of Fish and Wildlife. Institute for Natural Resources, Portland State University, Portland, OR, USA.

Hagmann, R. K., P. F. Hessburg, S. J. Prichard, N. A. Povak, P. M. Brown, P. Z. Fulé, R. E. Keane, E. E. Knapp, J. M. Lydersen, K. L. Metlen, M. J. Reilly, A. J. Sánchez Meador, S. L. Stephens, J. T. Stevens, A. H. Taylor, L. L. Yocom, M. A. Battaglia, D. J. Churchill, L. D. Daniels, D. A. Falk, P. Henson, J. D. Johnston, M. A. Krawchuk, C. R. Levine, G. W. Meigs, A. G. Merschel, M. P. North, H. D. Safford, T. W. Swetnam, and A. E. M. Waltz. 2021. Evidence for widespread changes in the structure, composition, and fire regimes of western North American forests. Ecological Applications 31(8):e02431. 10.1002/eap.2431