Late successional mixed conifer forests provide a multi-layered tree canopy, including large-diameter trees, shade-tolerant tree species in the understory, and a high volume of dead wood, such as snags and logs.
Ecoregional Characteristics
Blue Mountains
A mixture of conifer species is found throughout many forest sites in the Blue Mountains ecoregion. Mixed conifer forests can be divided into two subtypes based on temperature and moisture conditions. Douglas-fir and grand fir are the primary late successional tree species in the warmer drier climates of this ecoregion. Ponderosa pine and western larch may also be present. The cool mixed conifer type is indicated by the addition of more moisture-demanding and cold-tolerant species, such as subalpine fir and Engelmann spruce, at upper elevations or along streams where cold water-drainage and deep frost eliminate some species. The understory in this ecoregion generally includes huckleberry, serviceberry, oceanspray, snowberry, wild ginger, goldthread, starflower, bead lily, and oak fern.
Coast Range
Late successional mixed conifer forests in the Coast Range are generally dominated by two types of conifer trees: Sitka spruce and Douglas-fir. Sitka spruce forests occur within a narrow fog- and salt-influenced strip along the coast and extend into some valleys. Soils tend to be deep, acidic, and well-drained. Sitka spruce dominates the overstory, but western hemlock, western redcedar, Douglas-fir, big leaf maple, and red alder may be present. The lush understory has salmonberry, vine maple, salal, evergreen huckleberry, sword fern, deer fern, and a high diversity of mosses and lichens. Inland, Douglas-fir forests dominate. The understory of Douglas-fir forests includes shrub and forb species, such as vine maple, salal, sword fern, Cascade Oregon grape, western rhododendron, huckleberries, twinflower, vanilla leaf, and oxalis. Due to high precipitation in both Sitka spruce and Douglas-fir forests, fires are infrequent but do occur during hot, dry, east wind conditions after prolonged drought. When fires do occur, they are likely to be high severity, stand-replacing events. Other disturbances include small-scale windthrow events and floods driven by atmospheric river storms.
East Cascades
Late successional mixed conifer forests span the eastern slopes of the Cascade Mountains. This habitat contains a wide variety of tree species with Douglas-fir, grand fir, and western hemlock as the most common forest tree species that co-dominate most overstories. Several other conifers may also be present, including western redcedar, western white pine, western larch, ponderosa pine, and lodgepole pine. Undergrowth vegetation in the East Cascades ecoregion includes vine maple, Oregon grape, huckleberry, oxalis, thimbleberry, manzanita, ceanothus, and twinflower. Many sites once dominated by Douglas-fir and ponderosa pine (formerly maintained by wildfire) may now be dominated by grand fir (a fire sensitive, shade-tolerant species).
Klamath Mountains
Late successional mixed conifer forests in the Klamath Mountains ecoregion are characterized by high tree diversity. Douglas-fir is usually dominant. Depending on site characteristics, other canopy trees may include white fir, sugar pine, ponderosa pine, and incense cedar. Port-Orford cedar occurs on moist sites, such as riparian areas. Jeffrey pine and knobcone pine occur on serpentine soils. Broadleaf trees, such as tanoak, canyon live oak, golden chinquapin, big leaf and vine maple, and Pacific madrone, may occur in the subcanopy. A range of understory communities may be present, including those mostly dominated by shrubs, forbs, or grasses, or may be relatively open. However, with an increase in frequency of droughts, high instances of Douglas-fir mortality in the Klamath Mountains is quickly shifting the composition of these forests, creating conditions that result in catastrophic wildfires.
West Cascades
Late successional mixed conifer forests are found scattered throughout the West Cascades Ecoregion. While Douglas -fir dominates these forests, western hemlock is almost always co-dominant and usually dominates the understory. In the absence of stand replacing disturbance, Douglas-fir forests eventually convert to western hemlock. Other common trees include grand fir and western redcedar in the northern portion of the ecoregion, or incense cedar, sugar pine, white fir, and western redcedar in the southern portion of the ecoregion. The understory has shrub and forb species, such as vine maple, salal, sword fern, Cascade Oregon grape, western rhododendron, huckleberries, twinflower, vanilla leaf, and oxalis.
Limiting Factors and Recommended Approaches
Limiting Factor: Loss of Structural Habitat Elements
Where historical stands were perpetuated for 200 to more than 1,000 years, commercial forestlands are now commonly harvested every 60 years or less, which limits the maintenance and future recruitment of late-successional characteristics. In addition, the number of large-diameter snags and logs, which contribute to understory structure, has been reduced over time through wildfire and timber harvest.
Recommended Approach
Develop programs, incentives, and market-based approaches to encourage longer rotations and strategically located areas where large-diameter trees predominate. Where feasible, maintain structural elements, such as large-diameter tall trees, snags, and logs. Create snags from green trees or high-cut stumps where maintaining snags is not feasible or where snag management goals are not being met. Maintain old forest stand structures on private industrial forest lands. Provide technical assistance to landowners to leave large-diameter downed wood, green trees, or snags in the upland portion of harvested forests, as well as along riparian areas, to provide benefits for a diversity of wildlife and fish. Follow Oregon Forest Practices Act rules, which help to maintain a diversity of structural components on the landscape.
Limiting Factor: Loss of Late Successional Stand Size and Connectivity
Late successional forest stands have been greatly reduced in size and connectivity, particularly at lower elevations. This can impact species that are highly adapted to late successional conditions, require large tracts of intact habitats, and/or species that have limited ability to move over long distances to find new suitable areas. It also allows edge species to compete with those adapted to extensive interior forest habitat.
Recommended Approach
Maintain existing plans to protect and develop late successional habitat. Use active management to accelerate development of late successional structural characteristics in key areas to expand existing patches into larger areas; these will provide greater blocks of habitat for species with large area requirements or those that require interior forest habitat and are vulnerable to “edge effects”. Continue to carefully plan forest practices to maintain connectivity (KCI: Barriers to Animal Movement), particularly when species vulnerable to fragmentation are present. ODFW has mapped Priority Wildlife Connectivity Areas (PWCAs) to provide information on places across the landscape with the highest overall value for facilitating wildlife movement.
Seek opportunities to coordinate management of public and private lands (e.g., All-Lands Approach) whenever possible to address conservation needs. Use voluntary conservation tools, such as financial incentives and forest certification to achieve conservation goals on private lands. Work to maintain a diversity of forest types and ages to support wildlife habitat connectivity and ecosystem function at a landscape scale.
HABITAT CHANGE TRENDS ANALYSIS
Late successional mixed conifer forest habitats have been greatly reduced in size and connectivity by timber harvest. Following disturbance, such as timber harvest, regenerating conifer forests often succeed to deciduous or mixed deciduous-conifer forests without active management (e.g. replanting with Douglas-fir), especially on the west side of the state. These deciduous forests are primarily dominated by red alder and bigleaf maple. To investigate the transition from late successional mixed conifer forests to deciduous forest, the Institute of Natural Resources (INR) analyzed the change in total area of west-side deciduous and mixed conifer-deciduous forests between 1851 and 2016. The analysis showed a massive 243% increase in deciduous forests between 1851 and 1998, likely reflecting a shift towards earlier successional forests following logging.